Examples of research articles that were featured on issue covers.

Additional copies (as PDFs) of older papers etc. are being progressively added. Please contact me for copies of other publications, or check back again.

Peer reviewed works

(* = students, researchers & postdocs from Greenwood lab)

In review & In prep. 

  1. Padgett AB, Hyland EG, West* CK, Burgener LK, Greenwood DR, Basinger JF. In review, January 2023. Paleogene paleohydrology of Ellesmere and Axel Heiberg Islands (Arctic Canada) from palustrine carbonates. Paleoceanography and Paleoclimatology
  2. Patel* NU, Greenwood DR, McLachlan SMS, Galloway JM, Pospelova V. In prep. Palynology and palaeoecology across the Cretaceous-Paleogene boundary on the east coast of Vancouver Island, British Columbia, Canada. TBA
  3. West* CK, Greenwood DR, Reyes AV, Reinhardt L, Basinger JF. In prep. Photoperiod limits poleward range shifts of forest taxa during intervals of greenhouse climate. TBA


  1. Cramwinckel MJ, Burls NJ, Fahad AA, Knapp S, West* CK, Reichgelt T, Greenwood DR, Chan W-L, Donnadieu Y, Hutchinson D, DeBoer AM, Ladant J-B, Morozova P, Niezgodzki I, Knorr G, Steinig S, Zhang Z, Zhu J, Feng R, Lunt D, Abe-Ouchi A, Inglis G. 2023. Global and regional-scale hydrological response to early Eocene warmth. Paleoceanography and Paleoclimatology, 38(6): e2022PA004542 (DeepMIP special issue: Huber M & Lunt DI, Eds), doi: 10.1029/2022PA004542 Open Access
  2. Kalyniuk* JE, West* CK, Greenwood DR, Basinger JF, Brown CM. 2023. The Albian vegetation of Alberta as a food source for the nodosaurid dinosaur Borealopelta markmitchelli. Palaeogeography, Palaeoclimatology, Palaeoecology, 611: 111356, doi: 10.1016/j.palaeo.2022.111356 Open Access
  3. Mathewes RW, Greenwood DR, Reichgelt T. 2023. Plant megafossils, palynomorphs, and paleoenvironment from the late middle to late Eocene Burnaby Mountain Flora, Huntingdon Formation, British Columbia, Canada. International Journal of Plant Sciences, 184(3): 214–235, doi: 10.1086/724156 Open Access


  1. Greenwood DR. 2022. A fossil beech fern (cf. Phegopteris (C. Presl) Fée) from Driftwood Canyon Provincial Park, British Columbia. The Canadian Field-Naturalist, 136(3): 201–205. doi: 10.22621/cfn.v136i3.3001 Open Access
  2. Greenwood DR, Conran JG, West* CK. 2022. Palm fronds from western Canada are the northernmost palms from the Late Cretaceous of North America and may include the oldest Arecaceae. Review of Palaeobotany and Palynology, 301: 104641, doi: 10.1016/j.revpalbo.2022.104641 Open Access
  3. Lowe* AJ, Diefendorf AF, Schlanser K, Super J. West* CK, Greenwood DR. 2022. Dynamics of deposition and fossil preservation at the early Eocene Okanagan Highlands of British Columbia, Canada: insights from organic geochemistry. Palaios, 37 (5): 185–200, doi: 10.2110/palo.2021.007
  4. Patel* N, Greenwood DR, Galloway JM, Greenwood CL & Desautels* M. 2022. A reconstruction of the early Paleocene palaeovegetation of Turtle Mountain, southwestern Manitoba, Canada. Palynology, 46(4): 2064555, doi: 10.1080/01916122.2022.2064555 [free PDF here]
  5. Reichgelt T, Greenwood DR, Steinig S, Conran JG, Scriven LJ, Zhu J, Hutchinson DK, Lunt DJ. 2022. Plant proxy evidence for high rainfall and productivity in the Eocene of Australia. Paleoceanography and Paleoclimatology, 37(6): e2022PA004418 (DeepMIP special issue: Huber M & Lunt DJ, Eds), doi: 10.1029/2022PA004418.
  6. Sluiter IRK, Reichgelt T, Schultz N, Holdgate GR, Greenwood DR. 2022. A new perspective on Late Eocene and Oligocene vegetation and paleoclimates of South-eastern Australia. Palaeogeography, Palaeoclimatology, Palaeoecology, 596: 110985, doi: 10.1016/j.palaeo.2022.110985.
  7. Williams CJR, Lunt DJ, Salzmann U, Reichgelt T, Inglis G, Greenwood DR, Farnsworth AJ, Maidment RI, Wing-Le Chan, Donnadieu Y, Hutchinson DK, de Boer AM, Ladant J-B, Morozova PA, Niezgodzki I, Knorr G, Steinig S, Zhongshi Zhang, Jiang Zhu, Huber M, Otto-Bliesner BL. 2022. The African monsoon during the early Eocene from the DeepMIP simulations. Paleoceanography and Paleoclimatology, 37(5): e2022PA004419 (DeepMIP special issue: Huber M & Lunt DI, Eds), doi: 10.1029/2022PA004419. Open Access


  1. Huurdeman EP, Frieling J, Reichgelt T, Bijl PK, Bohaty SM, Holdgate G, Gallagher SJ, Peterse F, Greenwood DR, Pross J. 2021. Rapid expansion of meso-megathermal rain forests into the southern high latitudes at the onset of the Paleocene-Eocene Thermal Maximum. Geology 48 (1): 40–44, doi: 10.1130/G47343.1
  2. Pigg KB, DeVore ML, Greenwood DR, Sundue MA, Schwartzburd P, Basinger JF. 2021. Fossil Dennstaedtiaceae and Hymenophyllaceae from the Early Eocene of the Pacific Northwest. International Journal of Plant Sciences 182(9): 793–807, doi: 10.1086/715633
  3. Rozefelds AC, Stull GW, Hayes P, Greenwood DR. 2021. The fossil record of the Icacinaceae in Australia supports long standing palaeo-Antarctic rainforest connections in southern high latitudes. Historical Biology 33(11): 2854–2864, doi: 10.1080/08912963.2020.1832089 [free PDF here]
  4. Sudermann* M, Galloway JM, Greenwood DR, Reinhardt L, West* CK. 2021. Palynostratigraphy of the early Paleogene Margaret Formation at Stenkul Fiord, Ellesmere Island, Nunavut, Canada. Palynology 45(3): 459–476,  doi: 10.1080/01916122.2020.1861121 Open Access


  1. Brown CM, Greenwood DR, Kalyniuk* JE, Braman DR, Henderson DM, Greenwood* CL, Basinger JF. 2020. Dietary palaeoecology of an Early Cretaceous armoured dinosaur (Ornithischia; Nodosauridae) based on floral analysis of stomach contents. Royal Society Open Science 7 (6): 200305, doi: 10.1098/rsos.200305 Open Access
  2. Greenwood DR, Conran JG. 2020. Fossil coryphoid palms from the Eocene of Vancouver, British Columbia. International Journal of Plant Sciences 181 (2): 224–240, doi: 10.1086/706450
  3. Greenwood DR, West* CK, Basinger JF. 2020. The Miocene Red Lake macroflora of the Deadman River Formation (Chilcotin Group), Interior Plateau, British Columbia, Canada. Acta Palaeobotanica 60(2): 213–250. doi: 10.35535/acpa-2020-0011 Open Access
  4. Mathewes RW, Greenwood DR, Love RL. 2020. The Kanaka Creek fossil flora (Huntingdon Formation), British Columbia, Canada—paleoenvironment and evidence for Paleocene age using palynology and macroflora. Can. J. Earth Sci. 57(3): 348–365, doi: 10.1139/cjes-2018-0325
  5. Prader S, Kotthoff U, Greenwood DR, McCarthy FMG, Schmiedl G, Donders TH. Jan. 2020.  New Jersey’s paleoflora and eastern North American climate through Paleogene–Neogene warm phases. Review of Palaeobotany and Palynology 279: 104224 doi: 10.1016/j.revpalbo.2020.104224
  6. Schlanser K, Diefendorf AF, Greenwood DR, Mueller KE, West* CK, Lowe* AJ, Basinger JF, Currano ED, Flynn AG, Fricke HC, Meyer HW, Peppe DJ. 2020. On geologic timescales, plant carbon isotope fractionation responds to precipitation similarly to modern plants and has a small negative correlation with pCO2. Geochimica et Cosmochimica Acta 270: 264–281, doi: 10.1016/j.gca.2019.11.023
  7. Schlanser KM, Diefendorf AF, West* CK, Greenwood DR, Basinger JF, Meyer HW, Lowe* AJ, Naake H. 2020. Conifers are a major source of sedimentary leaf wax n-alkanes when dominant in the landscape: Case studies from the Paleogene. Organic Geochemistry 147: 104069 doi: 10.1016/j.orggeochem.2020.104069
  8. West* CK, Greenwood DR, Reichgelt T, Vachon* JM, Lowe* AJ, Sudermann* M, Basinger JF. 2020. Paleobotanical proxies for early Eocene climates and ecosystems in northern North America from mid to high latitudes. Climate of the Past 16: 1387–1410, doi: 10.5194/cp-2020-32 Open Access


  1. West* CK, Greenwood DR, Basinger JF. 2019. The late Paleocene to early Eocene Arctic megaflora of Ellesmere and Axel Heiberg islands, Nunavut, Canada. Palaeontographica, Abt. B, 300 (1–6): 47–163, doi: 10.1127/palb/2019/0066 [email for PDF]
  2. Willard DA, Donders TH, Reichgelt T, Greenwood DR, Sangiorgi F, Peterse F, Nierop K, Frieling J, Schouten S, Sluijs A. 2019. Arctic vegetation, temperature, and hydrology during Early Eocene transient global warming events. Global & Planetary Change, 178: 139–152, doi: 10.1016/j.gloplacha.2019.04.012 Open Access


  1. Lowe* AJ, Greenwood DR, West* CK, Galloway JM, Sudermann* M, Reichgelt T. 2018. Plant community ecology and climate on an upland volcanic landscape during the Early Eocene Climatic Optimum: McAbee fossil beds, British Columbia, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology, 511: 433–448, doi:10.1016/j.palaeo.2018.09.010
  2. Lowe* AJ, West* CK, Greenwood DR. 2018. Volcaniclastic lithostratigraphy and paleoenvironment of the lower Eocene McAbee fossil beds, Kamloops Group, British Columbia, Canada. Canadian Journal of Earth Sciences, 55(8): 923–934,  doi: 10.1139/cjes-2018-0019
  3. Reichgelt T, West* CK, Greenwood DR. 2018. The relation between global palm distribution and climate. Scientific Reports, 8(1): 4721, doi:10.1038/s***************-2, Open Access  [URL]


  1. Eberle JJ, Greenwood DR. 2017. An Eocene brontothere and tillodonts (Mammalia) from British Columbia, and their paleoenvironments. Canadian Journal of Earth Sciences, 54(9): 981−992, doi: 10.1139/cjes-2017-0061 Open Access. [URL]
  2. Gosse JC , Ballantyne A, Barker J, Csank A, Fletcher T, Grant G, Greenwood DR, McPhee R, Rybczynski N. 2017. PoLAR-FIT: Pliocene Landscapes and Arctic Remains, Frozen in Time. Geoscience Canada, 44: 47−54. doi: 10.12789/geocanj.2017.44.116 [paywall]
  3. Greenwood DR, Keefe* RL, Reichgelt T, Webb JA. 2017. Eocene paleobotanical altimetry of Victoria’s Eastern Uplands. Australian Journal of Earth Sciences, 64(5): 625−637, doi: 10.1080/08120099.2017.1318793. [free PDF here]
  4. Greenwood DR, West* CK. 2017. A fossil coryphoid palm from the Paleocene of western Canada. Review of Palaeobotany and Palynology, 239: 55−65, doi: 10.1016/j.revpalbo.2016.12.002
  5. Prader S, Kotthoff U, McCarthy FMG, Schmiedl G, Donders TH, Greenwood DR. 2017. Vegetation and climate development of the New Jersey hinterland during the late Middle Miocene (IODP Expedition 313 Site M0027). Palaeogeography, Palaeoclimatology, Palaeoecology, 485: 854–868, doi: 10.1016/j.palaeo.2017.07.028
  6. Prebble JG, Reichgelt T, Mildenhall DC, Greenwood DR, Raine JI, Kennedy EM, Seebeck HC. 2017. Terrestrial climate evolution in the Southwest Pacific over the past 30 million years. Earth and Planetary Science Letters, 459: 136–144, doi: 10.1016/j.epsl.2016.11.006
  7. Wolfe AP, Reyes AV, Royer DL, Greenwood DR, Doria G, Gagen MH, Siver PA,, Westgate JA. 2017. Middle Eocene CO2 and climate reconstructed from the sediment fill of a subarctic kimberlite maar. Geology, 45(7): 619–622, doi:10.1130/G39002.1 Open Access Gold.


  1. ♥Greenwood DR, Pigg KB, DeVore ML. 2016. Eocene paleontology and geology of western North America. Canadian Journal of Earth Sciences, 53(6): 543–547, doi:10.1139/cjes-2016-0043
  2. ♥Greenwood DR, Pigg KB, Basinger JF, DeVore ML. 2016. A review of paleobotanical studies of the Early Eocene Okanagan (Okanogan) Highlands floras of British Columbia, Canada and Washington, USA. Canadian Journal of Earth Sciences, 53(6): 548–564, doi: 10.1139/cjes-2015-0177
  3. ♥Gushulak* AC, West* CK, Greenwood DR. 2016. Paleoclimate and precipitation seasonality of the Early Eocene McAbee megaflora, Kamloops Group, British Columbia. Canadian Journal of Earth Sciences, 53(6): 591–604, doi: 10.1139/cjes-2015-0160
  4. ♥Mathewes RW, Greenwood DR, Archibald* SB. 2016. Paleoenvironment of the Quilchena flora, British Columbia, during the Early Eocene Climatic Optimum. Canadian Journal of Earth Sciences, 53(6): 574590, doi: 10.1139/cjes-2015-0163
  5. ♥Moss* PT, Smith* RY, Greenwood DR. 2016. A window into mid-latitudinal Early Eocene environmental variability: A high-resolution palynological analysis of the Falkland site, Okanagan Highlands, British Columbia, Canada. Canadian Journal of Earth Sciences, 53(6): 605–613, doi: 10.1139/cjes-2015-0167

♥ The 5 papers in CJES above are part of a themed special issue published June 2016 called ‘Eocene paleontology and geology of western North America’, where Greenwood, Pigg, and DeVore were the guest editors of the special issue.


  1. West* CK, Greenwood DR, Basinger JF. 2015. Was the Arctic Eocene ‘rainforest’ monsoonal? Estimates of seasonal precipitation from early Eocene megafloras from Ellesmere Island, Nunavut. Earth & Planetary Science Letters, 427: 18–30, doi: 10.1016/j.epsl.2015.06.036


  1. Archibald* SB, Makarkin VN, Greenwood DR, Gunnell GF. 2014. The red queen and court jester in green lacewing evolution: bat predation and global climate change. Palaios, 29(5): 185–191, doi: 10.2110/palo.2013.089. NB: Feature article and cover of issue 29(5).
  2. Archibald* SB, Morse G, Greenwood DR, Mathewes RW. 2014. Fossil palm beetles refine upland winter temperatures in the Early Eocene Climatic Optimum. Proceedings of the National Academy of Sciences, 111 (22): 8095–8100, doi: 10.1073/pnas.1323269111
  3. Eberle JJ, Rybczynski N, Greenwood DR. 2014. Early Eocene mammals from the Driftwood Creek beds, Driftwood Canyon Provincial Park, northern British Columbia. Journal of Vertebrate Paleontology, 34(4): 739–746, doi: 10.1080/02724634.2014.838175 NB: invited review article, feature article and cover of issue 34(4). [free PDF here]
  4. Eldrett JS, Greenwood DR, Polling M, Brinkhuis H, Sluijs A. 2014. A seasonality trigger for carbon injection at the Paleocene-Eocene thermal maximum. Climate of the Past, 10: 759–769, doi: 10.5194/cp-10-759-2014. Open Access (NB: amended from and replaces earlier draft in Clim. Past. Discussions in 2013)
  5. Fletcher TL, Greenwood DR, Moss PT, Salisbury SW. 2014. Paleoclimate of the Late Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, central-western Queensland, Australia: new observations based on CLAMP and bioclimatic analysis. Palaios, 29(3): 121−128, doi: 10.2110/palo.2013.080.
  6. Kotthoff U, Greenwood DR, McCarthy FMG, Müller-Navarra K, Prader S, Hesselbo SP. 2014. Late Eocene to middle Miocene (33 to 13 million years ago) vegetation and climate development on the North American Atlantic Coastal Plain (IODP Expedition 313, Site M0027). Climate of the Past, 10: 1523−1539,  doi:10.5194/cp-10-1523-2014 Open Access. (NB: amended from and replaces earlier draft in Clim. Past. Discussions in 2013).


  1. Archibald* SB, Greenwood DR, Mathewes RW. 2013. Seasonality, montane beta diversity, and Eocene insects: testing Janzen’s dispersal hypothesis in an equable world. Palaeogeography, Palaeoclimatology, Palaeoecology, 371: 1–8.
  2. Archibald* SB, Mathewes RW, Greenwood DR. 2013. The Eocene apex of Panorpoid scorpionfly Family diversity. Journal of Paleontology, 87(4): 677–695.
  3. Dillhoff R, Dillhoff T, Greenwood DR, DeVore M, and Pigg K. 2013. The Eocene Thomas Ranch flora, British Columbia. Botany, 91(8): 514–529, doi: 10.1139/cjb-2012-0313
  4. Eldrett JS, Greenwood DR, Polling M, Brinkhuis H, Sluijs A. 2013. A seasonality trigger for carbon injection at the Paleocene-Eocene thermal maximum. Climate of the Past Discussions, 9: 5837–5854. doi:10.5194/cpd-9-5837-2013 (see final version above published in Climate of the Past) OA
  5. Greenwood DR, Hill CR, Conran JG. 2013. Prumnopitys anglica sp. nov. (Podocarpaceae) leaves from the Eocene of England. Taxon, 62(3): 565–580.
  6. Kotthoff U, Greenwood DR, McCarthy FMG, Müller-Navarra K, Hesselbo SP. 2013. Vegetation and climate development on the North American Atlantic Coastal Plain from 33 to 13 million years ago (IODP Expedition 313). Climate of the Past Discussions, 9: 6551–6603. doi:10.5194/cpd-9-6551-2013 (see final version in Clim. Past)
  7. Reichgelt T, Kennedy EM, Mildenhall DC, Conran JG, Greenwood DR, Lee DE. 2013. Quantitative palaeoclimate estimates for Early Miocene southern New Zealand: evidence from Foulden Maar. Palaeogeography, Palaeoclimatology, Palaeoecology, 378: 36–44, doi: 10.1016/j.palaeo.2013.03.019


  1. Greenwood DR, Herold N, Huber M, Müller RD, Seton M. 2012. Forum Reply. Early to middle Miocene monsoon climate in Australia. Geology, 40 (6): e274. doi:10.1130/G33384Y.1 Open Access
  2. Eberle JJ, Greenwood DR. 2012. Life at the top of the greenhouse Eocene world – a review of the Eocene flora and vertebrate fauna from Canada’s High Arctic. GSA Bulletin, 124 (1/2): 3–23, doi: 10.1130/B30571.1 PDF NB: feature review article and cover (see top of this page).
  3. Pross J, Contreras L, Bijl PK, Greenwood DR, Bohaty SM, Schouten S, Bendle JA, Röhl U, Tauxe L, Raine JI, Huck CE, van de Flierdt T, Jamieson SSR, Stickley CE, van de Schootbrugge B, Escutia C, Brinkhuis H and IODP Expedition 318 Scientists. 2012. Persistent near-tropical warmth on the Antarctic continent during the early Eocene epoch. Nature, 488: 73–77. doi:10.1038/nature11300
  4. Smith* RY, Basinger JF, Greenwood DR. 2012. Early Eocene plant diversity and dynamics in the Falkland flora, Okanagan Highlands, British Columbia, Canada. Palaeobiodiversity and Palaeoenvironments, 92(3): 309–328.


  1. Archibald* SB, Greenwood DR, Smith* RY, Mathewes RW, Basinger JF. 2011. Great Canadian Lagerstätten 1. Early Eocene Lagerstätten of the Okanagan Highlands (British Columbia and Washington State). Geoscience Canada, 38 (4): 155–164NB: feature review article and cover (see top of this page).
  2. Archibald* SB, Johnson KR, Mathewes RW, Greenwood DR. 2011. Intercontinental dispersal of giant thermophilic ants across the Arctic during early Eocene hyperthermals. Proceedings of the Royal Society B, 278: 3679–3686. doi: 10.1098/rspb.2011.0729 PDF
  3. Barke J, Abels HA, Sangiorgi F, Greenwood DR, Sweet AR, Donders T, Reichart GJ, Lotter AF, Brinkhuis H. 2011. Orbitally forced Azolla blooms and Middle Eocene Arctic hydrology: Clues from palynology. Geology, 39(5): 427−430. doi: 10.1130/G31640.1 PDF download from journal website (subscription req’d).
  4. Herold N, Huber M, Greenwood DR, Müller RD, Seton M. 2011. Early to middle Miocene monsoon climate in Australia. Geology, 39(1): 3−6. doi: 10.1130/G31208.1 Open access.


  1. Archibald* SB, Bossert WH, Greenwood DR, Farrell BD. 2010. Seasonality, the latitudinal gradient of diversity, and Eocene insects. Paleobiology, 36(3): 374–398.doi: 10.1666/09021.1
  2. Ballantyne AP, Greenwood DR, Sinninghe Damsté JS, Csank AZ, Eberle JJ, Rybczynski N. 2010. Significantly higher Arctic surface temperatures during the Pliocene indicated by multiple independent proxies. Geology, 38(7): 603–606, doi:10.1130/G30815.1.
  3. Greenwood DR, Basinger JF, Smith* RY. 2010. How wet was the Arctic Eocene rainforest? Precipitation estimates from Arctic Paleogene macrofloras. Geology, 38(1): 15–18. doi: 10.1130/G30218.1
  4. Smith* RY, Greenwood DR, Basinger JF. 2010. Estimating paleoatmospheric pCO2 during the Early Eocene Climatic Optimum from stomatal frequency of Ginkgo, Okanagan Highlands, British Columbia, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology, 293(1-2): 120–131. doi:10.1016/j.palaeo.2010.05.006 PDF


  1. Eldrett JS, Greenwood DR, Harding IC, Huber M. 2009. Increased seasonality through the Eocene to Oligocene transition in northern high latitudes. Nature, 459: 969–974. doi:10.1038/nature08069 PDF [reprint requests should be directed to J.S.E.]
  2. Smith* RY, Basinger JF, Greenwood DR. 2009. Depositional setting, floristics and paleoenvironment of the Early Eocene Falkland site, a new fossil flora locality from the Okanagan Highlands, British Columbia. Canadian Journal of Earth Sciences, 46 (11): 811–822. doi:10.1139/E09-053 PDF
  3. Steart* DC, Greenwood DR, Boon PI. 2009. The chemical constraints upon leaf decay rates: Taphonomic implications among leaf species in Australian terrestrial and aquatic environments. Review of Palaeobotany and Palynology, 157: 358–374. doi:10.1016/j.revpalbo.2009.06.008


  1. Scarr* MJ, Greenwood DR, Scarpaci C. 2008. Intrinsic variation associated with stomatal frequency within Victorian specimens of Acacia melanoxylon, Acmena smithii and Eucalyptus obliqua. A preliminary study: implications for palaeoclimatological reconstructions. Quaternary Australasia, 25(2): 9–15. PDF
  2. Schouten S, Eldrett J, Greenwood DR, Harding I, Baas M, Sinninghe Damsté JS. 2008. Onset of long-term cooling of Greenland near the Eocene-Oligocene boundary as revealed by branched tetraether lipids. Geology, 36(2): 147–150. doi: 10.1130/G24332A.1


  1. Greenwood DR. 2007. North American Eocene Leaves and Climates: From Wolfe and Dilcher to Burnham and Wilf. In: Jarzen D, Retallack G, Jarzen S & Manchester S (Eds.) Advances in Mesozoic and Cenozoic Paleobotany: studies in celebration of David L. Dilcher and Jack A. Wolfe. Courier Forschungsinstitut Senckenberg, 258: 95–108. PDF
  2. Basinger JF, Greenwood DR, Wilson PG, Christophel DC. 2007. Fossil flowers and fruits of capsular Myrtaceae from the Eocene of South Australia. Canadian Journal of Botany, 85(2): 204–215. DOI: 10.1139/B07-001 Email me for a PDF.


  1. Steart* DC, Boon PI, Greenwood DR. 2006. Overland transport of leaves in two forest types in southern Victoria, Australia and its implications for palaeobotanical studies. Proceedings of the Royal Society of Victoria, 118(1): 65–74. PDF


  1. Archibald* SB, Greenwood DR. 2005.The Okanagan Highlands: Eocene biota,environments, and geological setting, southern British Columbia, Canada and northeastern Washington, USA. Canadian Journal of Earth Sciences 42(2): 111–114, doi: 10.1139/E05-012 PDF
    • NB: Introduction to a themed issue of CJES (v. 42.2) where Archibald & Greenwood were the guest editors of the whole issue.
  2. Greenwood DR. 2005. Leaf form and the reconstruction of past climates. New Phytologist, 166(2): 355–357. – invited contribution.
  3. Greenwood DR. 2005. Miocene wood from the Latrobe Valley coal measures, Victoria, Australia. Alcheringa, 29: 351–363. PDF
  4. Greenwood DR. 2005. Leaf margin analysis: taphonomic constraints. Palaios, 20(5): 498–505. DOI: 10.2110/palo.2004.P04-58
  5. Greenwood DR, Christophel DC. 2005. The origins and Tertiary history of Australian “Tropical” rainforests. Ch. 18, pp. 336−373, In, Bermingham, E., C. Dick, and C. Moritz (Eds), Tropical Rainforests: Past, Present, and Future. 1004 p. University of Chicago Press, Chicago.
  6. Greenwood DR, Archibald* SB, Mathewes RW, Moss* PT. 2005. Fossil biotas from the Okanagan Highlands, southern British Columbia and northeastern Washington State: climates and ecosystems across an Eocene landscape. Canadian Journal of Earth Sciences 42(2): 167–185. DOI: 10.1139/E04-100
  7. Moss* PT, Greenwood DR, Archibald* SB. 2005. Regional and local vegetation community dynamics of the Eocene Okanagan Highlands (British Columbia – Washington State) from palynology. Canadian Journal of Earth Sciences 42(2): 187–204. DOI: 10.1139/E04-095
  8. Steart* DC, Greenwood DR, Boon PI. 2005. Paleoecological implications of differential biomass and litter production in canopy trees in Australian Nothofagus and Eucalyptus forests. Palaios, 20(5): 452–462. DOI: 10.2110/palo.2004.P04-57


  1. Carpenter RA, Hill RS, Greenwood DR, Partridge AD, Banks* MA. 2004. No snow in the mountains: Early Eocene plant fossils from Hotham Heights, Victoria, Australia. Australian Journal of Botany  52(6): 685–718. DOI: 10.1071/BT04032
  2. Greenwood DR, Wilf, P, Wing SL, and Christophel DC. 2004. Paleotemperature estimates using leaf margin analysis: Is Australia different? Palaios, 19(2): 129–142.


  1. Gallagher SJ, Greenwood DR, Taylor D, Smith AJ, Wallace MW, Holdgate GR. 2003. The Pliocene climatic and environmental evolution of southeastern Australia: evidence from the marine and terrestrial realm. Palaeogeography, Palaeoclimatology, Palaeoecology, 193: 349–382. DOI: 10.1016/S0031-0182(03)00231-1 PDF
  2. Greenwood DR, Christophel DC, Scarr* M. 2003. Leaf stomatal frequency in the Australian tropical rainforest tree, Neolitsea dealbata (Lauraceae) as a proxy measure of pCO2. Palaeogeography, Palaeoclimatology, Palaeoecology, 196: 375–393. DOI: 10.1016/S0031-0182(03)00465-6
  3. Greenwood DR, Moss* PT, Rowett A., Vadala* AJ, Keefe* RL. 2003. Plant communities and climate change in southeastern Australia during the early Paleogene. pp. 365–380, In, Wing SL, Gingerich PD, Schmitz B, and Thomas E (eds.) Causes and Consequences of Globally Warm Climates in the Early Paleogene. Geological Society of America Special Paper 369. PDF


  1. Steart* DC, Boon PI, Greenwood DR, Diamond NT. 2002. Transport of leaf litter in upland streams of south-eastern Australian Eucalyptus and Nothofagus forests. Archiv für Hydrobiologie, 156(1): 43−61. DOI: 10.1127/0003-9136/2002/0156-0043 PDF


  1. Greenwood DR. 2001. Climate – wood and leaves. Section 4.3.6, pp. 480−483, In, Briggs DEG & Crowther PR (Eds) Palaeobiology II. Blackwell Scientific, London. ISBN 0 632 05147 7 doi: 10.1002/9780470999295.ch119 PDF
  2. Greenwood DR, Haines PW, Steart* DC. 2001. New Species of Banksieaeformis and a Banksia ‘Cone’ (Proteaceae) from the Tertiary of central Australia. Australian Systematic Botany, 14(6): 870−890. PDF
  3. Vadala* AJ, Greenwood DR. 2001. Australian Paleogene vegetation and environments: evidence for palaeo-Gondwanic elements in the fossil records of Lauraceae and Proteaceae. pp. 196−221, In, Metcalfe I, Smith JMB & Davidson I (eds.) Faunal and floral migrations and evolution in SE Asia-Australasia. Swets & Zeitlinger Publishers, Lisse. PDF


  1. Greenwood DR, Conran J. 2000. The Australian Cretaceous and Tertiary monocot fossil record. pp. 52–62, In, Wilson KL and Morrison DA (Eds.) Monocots: Systematics and Evolution. CSIRO Publishing, Melbourne. 738 pp. PDF
  2. Greenwood DR, Vadala* AJ, Banks* M. 2000. Climate change and vegetation responses during the Paleocene and Eocene in southeastern Australia. GFF, 122(1): 65−66, doi: 10.1080/11035890001221065. PDF
  3. Greenwood DR, Vadala* AJ, Douglas, J.G. 2000. Victorian Paleogene and Neogene macrofloras: a conspectus. Proceedings of the Royal Society of Victoria, 112(1): 65−92. PDF


  1. Wilf P, Wing SL, Greenwood DR, Greenwood CL. 1998. Using fossil leaves as paleoprecipitation indicators: An Eocene example: Comment and Reply. Geology, 29: 92.
  2. Wilf P, Wing SL, Greenwood DR, Greenwood CL. 1998. Using fossil leaves as paleoprecipitation indicators: An Eocene example. Geology, 26(3): 203-206.  doi:10.1130/0091-7613(1998)026<0203:UFLAPI>2.3.CO;2  PDF


  1. Greenwood DR. 1996. Eocene monsoon forests in central Australia? Australian Systematic Botany, 9(2): 95−112. PDF
  2. Wing SL, Greenwood D. 1996. Eocene continental climates and latitudinal gradients. Reply to Comment by Jordan, G.J. Geology, 24(11): 1054.


  1. Benbow M, Alley NF, Lindsay M, Greenwood DR. 1995. Geological history and palaeoclimate.  pp. 208−217, Ch. 10. Tertiary. In, Drexel JH. & Preiss WV (Eds) The geology of South Australia. vol. 2. The Phanerozoic. South Australian Geological Survey, Bulletin 54.  347 pp. + map. PDF
  2. Callen RA, Alley NF, Greenwood DR. 1995. Lake Eyre Basin.  pp. 188-194, Ch. 10. Tertiary. In, Drexel JH & Preiss WV (Eds) The geology of South Australia. vol. 2. The Phanerozoic. South Australian Geological Survey, Bulletin 54.  347 pp. + map. PDF
  3. Greenwood DR, Wing SL. 1995. Eocene continental climates and latitudinal gradients. Geology, 23(11): 1040−1048. doi:10.1130/0091-7613(1995)023<1044:ECCALT>2.3.CO;2 PDF


  1. Basinger JF, Greenwood DR, Sweda T. 1994. Early Tertiary vegetation of Arctic Canada and its relevance to palaeoclimatic interpretation. pp. 176−198, In, Boulter MC and Fisher HC (Eds.) Cenozoic Plants and Climates of the Arctic. NATO ASI Series, vol. 27 I. Springer-Verlag, Berlin. doi: 10.1007/978-3-642-79378-3_13 PDF
  2. Greenwood DR. 1994. Palaeobotanical Evidence for Tertiary Climates. Ch. 4, pp. 44−59, In, Hill RS (Ed.) History of the Australian Vegetation: Cretaceous to Recent. Cambridge University Press, Cambridge. 433 pp. PDF
  3. Greenwood DR, Basinger JF. 1994. The paleoecology of high latitude Eocene swamp forests from Axel Heiberg Island, Canadian High Arctic. Review of Palaeobotany and Palynology, 81: 83−97. PDF


  1. Greenwood DR, Basinger JF. 1993. Stratigraphy and floristics of peat-coal layers of Eocene swamp forests from Axel Heiberg Island, Canadian Arctic Archipelago. Canadian Journal of Earth Sciences, 31: 1914−1923. PDF
  2. Greenwood DR, Behrensmeyer AK, Wing SL. 1993. Determining Paleoclimates: Evolution of Atmospheres, Is the Geological Past a Key to the (Near) Future? Science, 260: 278−279. PDF
  3. Wing SL, Greenwood DR. 1993. Fossils and fossil climates: the case for equable Eocene continental interiors. Philosophical Transactions of the Royal Society, London B, 341: 243−252. PDF
  4. Wing SL., Greenwood DR. 1993. Fossils and fossil climates: the case for equable Eocene continental interiors. Ch. 5, pp. 243−252, In, Allen JRL, Hoskins BJ, Sellwood BW & Spicer RA (Eds), Palaeoclimates and their modelling with special reference to the Mesozoic Era. Chapman & Hall, London. (also published as Phil. Trans. Roy. Soc. London B. Biol. Sci., 341 as indicated above)


  1. Christophel DC, Scriven LJ, Greenwood DR. 1992. An Eocene megaflora from Nelly Creek, South Australia. Transactions of the Royal Society of South Australia, 116(2): 65−76. PDF
  2. Greenwood DR. 1992. Taphonomic constraints on foliar physiognomic interpretations of Late Cretaceous and Tertiary palaeoclimates. Review of Palaeobotany and Palynology, 71: 142−196. PDF


  1. Christophel DC, Greenwood DR. 1989. Changes in climate and vegetation in Australia during the Tertiary. Review of Palaeobotany and Palynology, 58: 95−109. PDF
  2. Christophel DC, Greenwood DR. 1988. A comparison of Australian tropical rainforest and Tertiary fossil leaf-beds. Proceedings of the Ecological Society of Australia, 15: 139−148. PDF
  3. Greenwood DR. 1991. The Taphonomy of Plant Macrofossils. Ch. 7, pp. 141−169, In, Donovan SK. (Ed.) The Processes of Fossilization. Belhaven Press, London, 303 pp. PDF


  1. Christophel DC, Greenwood DR. 1987. A megafossil flora from the Eocene of Golden Grove, South Australia. Transactions of the Royal Society of South Australia, 111(3): 155−162. PDF
  2. Greenwood DR. 1987. Early Tertiary Podocarpaceae: megafossils from the Eocene Anglesea locality, Victoria, Australia. Australian Journal of Botany, 35(2): 111−134. PDF 

Comments, Unrefereed papers, extended abstracts, ‘letters’, reviews, and internal reports.


  1. Kalyniuk, JE, Greenwood, DR, & Greenwood CL. 2022. An armoured dinosaur’s
    last supper. Research Connection vol. 3. Passion and enthusiasm drive discovery. pp. 40-41.
  2. Greenwood, D & West, C. 2022. When Canada had palm tree fringed tropical beaches. Research Connection, 4(2): 2 pp.


  1. Kalyniuk, JE, Greenwood, DR, & Greenwood, CL. 2020. An armored dinosaur’s last supper. Research Connection, 1(3); 2 pp.


  1. Sudermann, M & Greenwood, D. 2019. Can ancient Arctic swamps help us predict the effects of climate change? Research Connection, 3(2): 2 pp.


  1. Lowe A & Greenwood D. 2017. Fossil plants and global warming. Research Connection, 2(3): 2 pp.


  1. Prader, S., Kotthoff, U., McCarthy, F. and Greenwood, D. Vegetation and climate development on the Atlantic Coastal Plain during the late Mid-Miocene Climatic Optimum (IODP Expedition 313). Geophysical Research Abstracts Vol. 17, EGU2015-10866, 2015.


  1. Greenwood, D.R. Conference report: Geological Society of America Annual Meeting 2014 Vancouver, British Columbia (Topical Session T201. Eocene Northern North America: Biotic Change and Environmental Context). Geolog Vol. 43 (4): 24−25.


  1. Bruce, G. 2013. Interview with D. Greenwood. A warmer Brandon will see more storms. Brandon Sun, July 20th, 2013.


  1. Rombough, P. and Greenwood, D.R. Deniers provide ‘facts’ with no bite. Letters to the Editor, Brandon Sun, December 19, 2006. PDF
  2. Blair, D., Greenwood, D.R. and Hanesiak, J. 2006. Global threat. Awash in evidence, global warming deniers continue to deceive the public. View from the West. Winnipeg Free Press, May 18, 2006. p. A14.


  1. Greenwood, D.R. 2005. Special dedication volume. IOP Newsletter 79: 5.
  2. Greenwood, D.R. 2005. Research Reports. Victoria University. Nomen Nudum 29: 85-87. (AAP) (ISSN 1447-4662)
  3. Haggart. J.W., Archibald, S.B., Basinger, J.F., Greenwood, D.R., Mathewes, R.W. and Wilson, M.V.H. 2005.  Significance of the Horsefly Fossil Site, British Columbia. British Columbia Paleontological Alliance. PDF
  4. Archibald, S.B., Pigg, K.P., Greenwood, D.R., Manchester, S.R., Barksdale, L., Johnson, K.R., Sternberg, M., Stockey, R.A., DeVore, M.L., and Rothwell, G.W. 2005. Wes Wehr dedication. Canadian Journal of Earth Sciences 42(2): 115−117. [PDF available from Journal website]


  1. Greenwood, D.R. 2003. Reports of Meetings. Climate and Biota of the Paleogene, Northwest College, Powell, Wyoming, USA. Nomen Nudum 28: 43-45. (Association of Australasian Palaeontologists) (ISSN 1447-4662)
  2. Greenwood, D.R. 2003. Research Reports. Victoria University. Nomen Nudum 28: 96-97. (AAP) (ISSN 1447-4662)


  1. Greenwood, D. 2000. Conference review. Palaeobotanists meet in Qinhuangdao, P.R. China. Australian Systematic Botany Society Newsletter, 104: 22−23. ISSN 1034-1218
  2. Greenwood, D.R. 2000. Australia’s cycad fossils and the antiquity of the Daintree rainforest. Palms and Cycads (ISSN 1321-2346), 66: 2−9. [web version]


  1. Greenwood, D.R., Vadala, A., & Banks, M. 1999. Climates and changes in forest floristics during the Paleocene and Eocene in southeastern Australia. In, Andreasson, F.P., Schmitz, B. & Thompson, E.I. (Eds.), Early Paleogene warm climates and biosphere dynamics. Abstract Volume. ISSN 1400-383X. Göteborg University, Sweden. (also listed below)
  2. Latham, S. & Greenwood, D. 1999. Innovative geography at the tertiary level. Interaction (Journal of the Geog. Teachers’ Assoc. Vic. Inc.), 27(4): 12−14. – invited contribution.
  3. Greenwood, D.R. 1999. Tertiary palaeobotany in Melbourne. IOP Newsletter, 67: 7.
  4. Greenwood, D.R. 1998. Palaeobotany in Melbourne. IOP Newsletter, 65: 5-6.
  5. Greenwood, D.R. 1998. Life in the greenhouse. Nexus: News from Victoria University, 8(13): 4.


  1. Greenwood, D.R. 1996. Palaeobotany on the INTERNET. Palaeoaustral (Newsletter of the Palynological & Palaeobotanical Assoc. of Australasia ISSN 1321-9677), Number 4, pg. 3.


  1. Greenwood, D.R. 1994. An Australian Institute of Biological Sciences, ASBS, and large conferences. Australian Systematic Botany Society Newsletter, 81: 11−12.
  2. Greenwood, D.R. 1994. Report on the nature of plant detritus found blocking the drain of the Adelaide Botanic Gardens Main Lake, following the Dec. 13 1993 flooding of the University of Adelaide Medical School. unpubl. report prepared for Thompsons, Solicitors & Barristers, Adelaide. 12 pp.
  3. Greenwood, D.R. 1994. Palaeobotany. Australian Systematic Botany Society Newsletter, 81: 22−23.

pre 1994

  1. Greenwood, D.R. & Collinson, M.E. 1992. The origins and Early Tertiary history of modern plant communities. pg. 113, In, Lidgard, S. & Crane, P.R. (eds), Fifth North American Paleontology Convention. Abstracts and Program. Paleontology Society Special Publ. 6. 329 pp.
  2. Greenwood, D.R. 1991. Middle Eocene megafloras from central Australia: earliest evidence for Australian sclerophyllous vegetation. Supplement to American Journal of Botany (Abstracts), 78(6): 114 – 115.
  3. Greenwood, D.R., Callen, R.A. & Alley, N.F. 1990. The correlation and depositional environment of Tertiary strata based on macro-floras in the Southern Lake Eyre Basin. South Australian Dept. Mines & Energy. Report Book No. 90/15. 57 pp. & 13 figs.
  4. Greenwood, D. 1990. Museum cuts. New Scientist, 126 (1722): 75-76.
  5. Greenwood, D.R. 1990. Fijian Rainforest Trees. 2 pp. Philatelic Bulletin No. 98. Philatelic Bureau, Fiji Post & Telecommunications, Suva.
  6. Greenwood, D.R. 1988. A Field Guide to the Eocene Anglesea Locality. 3rd Conference of the International Organisation of Palaeobotany, Melbourne, 1988. 14 pp. University of Adelaide.